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Coyote in the wild. (Photo by Paul Rego) |
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At my office building in the back of a room relegated for storage, sit cardboard boxes with contents from the days of carbon paper and manual typewriters including old memos, articles and photos. One black and white photo shows a man displaying a |
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small, dead coyote laid on the snow. I imagine that the photo dates to the 1950s when the first coyotes were appearing in Connecticut and the uniqueness of the odd, never before seen specimen drew special attention, perhaps to the point of deserving a photo. Now, some four decades later, Connecticut seems a landscape saturated with coyotes. This adaptable species is found in remote forests and residential suburbs, from farm fields to golf courses. Coyotes have dramatically spread across North America. Until about one-hundred years ago the species only ranged through the western states -- out with the antelope, cowboys, and howl-inducing prairie moons: coyotes are integral to western folklore. Then like a prairie wildfire the population began to spread past former boundaries, extending north into Canada, south into Mexico and, as if assisted by prevailing westerlies, east across the Mississippi River through the upper Midwest and southern Canada. One could be convinced that New England was the goal of this latter, eastward vector, |
Photo
of dead coyote found in Connecticut in the
1950's. |
Many factors may have contributed to the coyote's rapid range expansion. One plausible theory holds that coyotes were filling a void or open niche created by the extirpation of wolves and mountain lions. Another factor may have been the increase in some prey populations, especially deer, during the same time as the expansion. Theories aside, coyotes are biologically well equipped to pioneer new territory. They are adaptable in diet and habitat use, fecund, and mobile. And, contrary to the rumor that periodically surfaces, the government did not introduce coyotes into the east to control burgeoning deer populations. (Some coyotes were privately stocked in southeastern states and this may have contributed to their presence there.)
Several studies in the northeast have investigated the biology of this recently arrived predator. Coyotes in the east differ from western coyotes, enough so that some feel they deserve distinct subspecies status. The most obvious difference is size -- eastern coyotes are an average of five to ten pounds heavier than western coyotes. In Connecticut adult females weigh 30 to 35 pounds and adult males weigh 35 to 40 pounds. Rarely do they exceed 50 pounds, although claims of 70 and 80 pounders can be heard atmany coffee shops (or other beverage-based gatherings). As often as not, the vernacular moniker "coydog" accompanies these claims. The best evidence indicates this is a misnomer -- Canis familiaris (the common dog)is not in the eastern coyote's pedigree,
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but, Canis lupus (the timber wolf) is. A small amount of wolf lineage, probably a result of contact between the two species during the eastward expansion through Canada, may account for the larger size in the east. Another difference is the wide range of coloration seen in eastern coyotes; they vary from blonde to red to charcoal gray. The standard coloration is a light brown with dark highlights concentrated on the back and light cream on the underparts. Standard equipment that distinguishes coyotes from other canines includes erect ears; a relatively narrow, pointed muzzle; and a full, bottle-brush tail that is held straight and lowered. |
They are both predator and scavenger, particularly when the food is venison. In winter, when alternative foods such as small mammals and fruits become scarce, remains of white-tailed deer are common in stomachs and scats. Discerning whether the deer was killed, a feat within a coyote's capabilities, or scavenged is difficult from this evidence. Birds, both wild and domestic, are eaten. Coyotes can attack livestock and pets as well. Each year about one dozen attacks on sheep are reported in Connecticut. Reports of attacks on cats are common, while reported dog attacks number about three to six per year. Coyotes are not known for raiding garbage cans but will selectively eat table scraps left in backyard compost piles. Perhaps the most bizarre gastronomical account reported to me involved coyotes visiting a backyard to dine on dog droppings. In short, the diet is diverse and opportunistic.
Several studies have also shed light on the eastern coyote's social organization. Researchers in Maine and Vermont captured coyotes with leg-hold traps (contrary to popular perception these traps, especially the padded versions, can capture animals with minimal injury) and fit them with radio collars. Radio tracking revealed that the social grouping of coyotes is centered around an adult pair. Their mating in February results in an April litter. Research in Connecticut has estimated the litter size at birth to be seven, and found that nearly all adult females breed. The pups are raised by both adults and remain with them until late fall and winter arrive. Nearly adult size, near the summit of independence, most disperse from the adults' home range. As in many wild animals, mortality strikes heavy on the young and perhaps only three to five young survive to independence. The adult pair remain in the home range and the cycle is repeated.
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Each family group's home range is exclusive, like pancakes on a griddle, with little or no overlap with neighboring home ranges. In Connecticut these "pancakes" probably encompass ten square miles. With such a high reproductive rate and three to four decades to establish home ranges across the terrestrial "griddle" that is Connecticut, the nutmeg coyote population has probably reached its upper limit of growth, and is estimated to number from three to five thousand. What is the fate of dispersing young now that Connecticut is fully inhabited by coyotes? They can travel remarkable distances, some have traveled 200 miles, but with few vacant home ranges most probably do not survive. What are the consequences of cohabitating with this nonnative canid? The ecological consequences are not black and white, quantified and defined as many people wish, rather they are complex and dynamic, occurring in shades of gray. One consequence, revealed by several telemetry studies, is that red foxes appear to be excluded from coyote home ranges, therefore the advent of coyotes has been negative for fox populations. Although coyotes kill deer, we believe that their territoriality and the high deer densities at our latitude will preclude them from controlling deer populations. Relationships with other wildlife are not well studied. Turkey hunters that wonder if coyotes will decimate the turkey |
The consequences from a human perspective are varied. Farmers (both hobby and vocational) must contend with a new threat to livestock. Birdwatchers, hunters, and hikers have a new animal to accent outings, both visually and audibly. A small number of hunters have a new, exceptionally challenging game species. Airport managers have another runway safety hazard. (Coyotes find the open fields of airports quite attractive.) Pet owners have a new consideration in caring for their outdoor pets. Some have had